1. U.S. Geological Survey Utah Cooperative Fish and Wildlife Research Unit, Department of Wildland Resources, Utah State University, Logan, UT, USA
2. Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT, USA
3. Center for Biodiversity and Global Change, Yale University, New Haven, CT, USA
4. Department of Fish and Wildlife Conservation, Virginia Tech, 310 West Campus Dr., Blacksburg, VA, USA