1. Christ’s College, University of Cambridge , Cambridge CB2 3BU , UK
2. Department of Zoology, University of Cambridge , Cambridge CB2 3EJ , UK
3. School of Biosciences, Cardiff University , Cardiff CF10 3AX , UK
4. School of Natural Sciences, Bangor University , Bangor, Gwynedd LL57 2UW , UK
5. Aquatic Ecology Unit, Department of Biology, Lund University , Lund , Sweden