1. Ethology lab, Ethology Ecology and Evolution group, Cavanilles Institute of Biodiversity and Evolutionary Biology, University of Valencia , C/Catedrático José Beltrán 2, 46980, Valencia , Spain
2. Department of Biology, Lund University , 221 00 Lund , Sweden
3. School of Biological Sciences, University of East Anglia , Norwich, NR4 7TJ , UK
4. Division of Ecology and Evolution, Research School of Biology, Australian National University , Canberra ACT 2600 , Australia