1. State Key Laboratory of Integrated Management of Pest Insects and Rodents, Institute of Zoology, Chinese Academy of Sciences , Beijing 100101 , China
2. College of Life Science, University of Chinese Academy of Sciences , Beijing 100049 , China
3. Centre for Evolutionary Biology, School of Biological Sciences (M092), University of Western Australia , 35 Stirling Hwy, Crawley, WA 6009 , Australia
4. Ministry of Health Institute of Laboratory Animal Science, Chinese Academy of Medical Sciences and Comparative Medicine Centre, Peking Union Medical College , Beijing 100730 , China
5. CAS Center for Excellence in Biotic Interactions, University of Chinese Academy of Sciences , Beijing 100049 , China