1. Department of Ecology and Genetics, University of Oulu, Pentti Kaiteran katu, Oulu, Finland
2. Department of Environmental Science, Policy, and Management, University of California, Berkeley, Berkeley, CA, USA
3. Aquatic population dynamics, Natural Resources Institute Finland (Luke), Paavo Havaksen tie, Oulu, Finland
4. Department of Environmental and Biological Sciences, University of Eastern Finland, Joensuu, Finland
5. Aquatic population dynamics, Kainuu Fisheries Research Station, Natural Resources Institute Finland (Luke), Manamansalontie,Paltamo, Finland