1. Hebei Normal University of Science and Technology Hebei Key Laboratory of Horticultural Germplasm Excavation and Innovative Utilization, College of Horticulture Science and Technology, , Qinhuangdao 066004, China
2. China Agricultural University State Key Laboratories of Agrobiotechnology, Joint International Research Laboratory of Crop Molecular Breeding, Beijing Key Laboratory of Growth and Developmental Regulation for Protected Vegetable Crops, Department of Vegetable Sciences, , Beijing 100193, China
3. Qingdao Agricultural University Engineering Laboratory of Genetic Improvement of Horticultural Crops of Shandong Province, College of Horticulture, , Qingdao 266109, China