1. Department of Animal Biosciences, University of Guelph , Guelph, ON N1G 2W1 , Canada
2. Agriculture and Agri-Food Canada, Sherbrooke R & D Centre, 2000 College , Sherbrooke, QC J1M 0C8 , Canada
3. Department of Population Medicine, University of Guelph , Guelph, ON N1G 2W1 , Canada