1. Department of Biological Sciences, Graduate School of Humanities and Sciences, Ochanomizu University , Ohtsuka, Bunkyo-ku, Tokyo 112-8610, Japan
2. RIKEN Center for Brain Science-Evident Open Collaboration Center, Center for Brain Science (CBS) , RIKEN, 2-1, Hirosawa, Wako-shi, Saitama 351-0106, Japan