1. Department of Biomedical Informatics, Colorado Center for Personalized Medicine, University of Colorado Anschutz Medical Campus , Aurora, Colorado , USA
2. Center for Human Genetics and Department of Genetics and Biochemistry, Clemson University , South Carolina , USA
3. Precision Medicine Institute, Macrogen Inc. , Seoul , Republic of Korea
4. Department of Anthropology and Carl R. Woese Institute for Genomic Biology, University of Illinois Urbana-Champaign, Illinois , USA
5. Department of Ecology, Evolution, and Organismal Biology and Center for Computational and Molecular Biology, Brown University , Providence, Rhode Island , USA
6. Institute at Brown for Environment and Society, Brown University , Providence, Rhode Island , USA
7. Department of Anthropology, University of Colorado Boulder , Colorado , USA