1. Department of Biological Sciences and the director of the Genetic and Genomic Scholars graduate program, North Carolina State University, Raleigh, North Carolina, United States
2. Department of Biological Sciences and director of Herbarium UTEP, University of Texas, El Paso, El Paso, Texas, United States
3. University of Connecticut, Mansfield, Connecticut, United States, and editor-in-chief of The American Naturalist, Chicago, Illinois, United States
4. Utah State University, Logan, Utah, United States
5. University of Texas at Austin, Austin, Texas, United States, The author order was determined by a random number generator