1. Max Planck Institute for Animal Behavior, Bodensee, Germany
2. Institute for Communities and Wildlife, Africa, Department of Biological Sciences, University of Cape Town, Cape Town, South Africa
3. Behavioural Ecology and Endocrinology Laboratory and Andrew King is an associate professor and head of the SHOAL group in the Department of Biosciences at Swansea University, Swansea, Wales, United Kingdom