1. College of Life and Environment Sciences, Minzu University of China , Beijing 100081 , China
2. Key Laboratory of Animal Ecology and Conservational Biology, Institute of Zoology, Chinese Academy of Sciences , Beijing 100101 , China
3. College of Wildlife and Protected Area, Northeast Forestry University , Harbin 150040 , China
4. College of Resources and Environmental Sciences, Nanjing Agricultural University , Nanjing 210095 , China