1. Division of Gastroenterology, Department of Medicine, University of Michigan, AnnArbor, MI, USA
2. University of Michigan Crohn’s and Colitis Program, University of Michigan, AnnArbor, MI, USA
3. Tokyo R&D Center, Miyarisan Pharmaceutical, Tokyo, Japan
4. Department of Surgery, University of Michigan, AnnArbor, MI, USA
5. Department of Cell and Developmental Biology, University of Michigan, AnnArbor, MI, US