1. Centre of Excellence for Gastrointestinal Inflammation and Immunity Research, University of Alberta , Edmonton, AB, T6G 2X8 , Canada
2. Department of Physiology, University of Alberta , Edmonton, AB, T6G 1C9 , Canada
3. Department of Pediatrics, University of Alberta, Edmonton Clinic Health Academy , Room 4-577, 11405 87th Ave., Edmonton, AB. T6G 1C9 , Canada