1. Department of Botany and Plant Pathology, Purdue University, West Lafayette, IN, USA
2. Purdue Center for Plant Biology, Purdue University, West Lafayette, IN, USA
3. Center of Pear Engineering Technology Research, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Nanjing Agricultural University, Nanjing, Jiangsu, China
4. Division of Biology and Biological Engineering, California Institute of Technology, Pasadena, CA, USA
5. Howard Hughes Medical Institute, California Institute of Technology, Pasadena, CA, USA