1. School of Horticulture, Hainan University, Haikou, China
2. Key Laboratory for Quality Regulation of Tropical Horticultural Crops of Hainan Province, Haikou, China
3. Innovation Cluster of Crop Molecular Biology and Breeding, Anhui Agricultural University, Hefei, China
4. School of Agronomy, Anhui Agricultural University, Hefei, China
5. School of Environmental and Life Sciences, The University of Newcastle, Callaghan, NSW, Australia