1. Division of Forest and Biomaterials Science, Graduate School of Agriculture, Kyoto University, Kyoto, Japan
2. Department of Zoology, Graduate School of Science, Kyoto University, Kyoto, Japan
3. Institute of Natural and Environmental Sciences, University of Hyogo, Sanda, Japan
4. Graduate School of Humanities and Social Sciences, Hiroshima University, Higashi-Hiroshima, Japan