1. Hachinohe Laboratory, Tohoku National Fisheries Research Institute, Japan Fisheries Research and Education Agency, Shimomekurakubo, Same, Hachinohe, Aomori, Japan
2. Tohoku National Fisheries Research Institute, Japan Fisheries Research and Education Agency, Shinhama, Shiogama, Miyagi, Japan
3. National Research Institute of Fisheries Science, Japan Fisheries Research and Education Agency, Fukuura, Yokohama, Kanagawa, Japan