1. Centre de Recherche en Paléontologie – Paris (CR2P, UMR 7207), (CNRS, MNHN, Sorbonne Université), Muséum national d’Histoire naturelle, case postale 38, 57 rue Cuvier, F-75231 Paris cedex 05, France
2. Hokkaido Research Organization, Central Fisheries Research Institute, 238 Hamanaka, Yoichi, 045-8555 Hokkaido, Japan
3. State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China