1. Kansas Biological Survey, and the Natural History Museum (Biodiversity Institute), Kansas University, Lawrence, KS, USA
2. South African Environmental Observation Network, P.O. Box 110040, Hadison Park, Kimberley, 8306, Republic of South Africa
3. Centre for Environmental Management, University of the Free State, P.O. Box 339, Bloemfontein, 9300Republic of South Africa