1. Horseshoe Crab Research Group, Faculty of Science and Marine Environment, Universiti Malaysia Terengganu, Kuala Nerus, Malaysia
2. Horseshoe Crab Research Group, Institute of Marine Biotechnology, Universiti Malaysia Terengganu, Kuala Nerus, Malaysia
3. Center for Ecosystem Management and Natural Resources, Faculty of Science and Technology, Universiti Kebangsaan, Selangor, Malaysia
4. Institute of Tropical Biodiversity and Sustainable Development, Universiti Malaysia Terengganu, Kuala Nerus, Malaysia