1. Graduate School of Life Sciences, Tohoku University , 2-1-1 Katahira, Aoba-ku, Sendai, Miyagi 980-8577, Japan
2. PRESTO, Japan Science and Technology Agency (JST) , 4-1-8, Honcho, Kawaguchi, Saitama 332-0012, Japan
3. Plant Genetics, TUM School of Life Sciences, Technical University of Munich (TUM) , Emil Ramann Str. 4, 85354 Freising, Germany