1. College of Life Sciences, University of Chinese Academy of Sciences , 19A Yuquan Rd., Shijingshan District, Beijing 100049, China
2. State Key Laboratory of Protein and Plant Gene Research, Peking-Tsinghua Center for Life Sciences, School of Life Sciences, Center for Quantitative Biology, Peking University , 5 Summer Palace Rd., Haidian District, Beijing 100871, China