1. RIKEN Center for Sustainable Resource Science, 1-7-22 Suehiro-cho, Tsurumi, Yokohama, Kanagawa 230-0045, Japan
2. Department of Biological Sciences, Faculty of Science, The University of Tokyo, 7-3-1 Hongo, Bunkyō-ku, Tōkyō 113-8654, Japan
3. Institut de Biologie Moléculaire des Plantes, Université de Strasbourg, 12 rue du Général Zimmer, Strasbourg 67084, France
4. Department of Biology, Faculty of Science, Niigata University, Ikarashi, Niigata 950-2181, Japan
5. College of Bioscience and Biotechnology, Chubu University, 1200 Matsumoto-cho, Kasugai, Aichi 487-8501, Japan
6. UMR5004 Biochimie et Physiologie Moléculaire des Plantes, Université de Montpellier, CNRS, INRAE, Institut Agro, 2 place Pierre Viala, Montpellier 34060, France
7. Leibniz-Institut für Gemüse- und Zierpflanzenbau (IGZ) e.V., Theodor-Echtermeyer-Weg 1, Großbeeren 14979, Germany