1. Department of Biology, Graduate School of Science, Kobe University , 1-1 Rokkodai, Kobe, 657-8501 Japan
2. Department of Biological Sciences, Graduate School of Science, The University of Tokyo , 7-3-1 Hongo, Bunkyo-Ku, Tokyo, 113-0033 Japan
3. College of Life Sciences, Ritsumeikan University , 1-1-1 Noji-higashi, Kusatsu, 525-8577 Japan