1. Graduate School of Agricultural and Life Sciences, The University of Tokyo , 1-1-1 Yayoi, Bunkyo-ku, Tokyo, 113-8657 Japan
2. Research Center for Agricultural Information Technology, National Agriculture and Food Research Organization, Kouwa Nishi-Shimbashi Bldg . 5f, 2-14-1 Nishi-Shimbashi, Minato-ku, Tokyo 105-0003, Japan
3. Department of Life Sciences, Faculty of Agriculture, Meiji University , 1-1-1 Higashi-Mita, Tama-ku, Kawasaki, Kanagawa, 214-8571 Japan
4. Institute of Plant Science and Resources, Okayama University , 2-20-1 Chuo, Kurashiki, Okayama, 710-0046 Japan