1. Graduate School of Biostudies, Kyoto University , Kyoto, 606-8502 Japan
2. Department of Plant Sciences, University of Cambridge , Cambridge CB2 3EA, UK
3. National Institute of Genetics, Research Organization of Information and Systems , Mishima, 411-8540 Japan
4. Faculty of Science and Technology, Tokyo University of Science , Noda, 278-8510 Japan
5. Faculty of Biology-Oriented Science and Technology (BOST), Kindai University , Kinokawa, 649-6493 Japan
6. School of Biological Sciences, Monash University , Melbourne 3800, Australia
7. ARC Centre of Excellence for Plant Success in Nature and Agriculture, Monash University , Melbourne 3800, Australia