1. Johns Hopkins Center on Aging and Immune Remodeling and Division of Geriatric Medicine and Gerontology, Department of Medicine, Johns Hopkins University School of Medicine , Baltimore, MD 21224 , United States
2. Department of Immunology, University of Tübingen , Tübingen 72076 , Germany
3. Health Sciences North Research Institute , Sudbury, ON P3E 2H2 , Canada