1. School of GeoSciences, Grant Institute, The King’s Buildings, University of Edinburgh , James Hutton Road, Edinburgh, EH9 3FE , UK
2. Department of Natural Sciences, National Museum of Scotland , Edinburgh, EH1 1JF , UK
3. Department of Biomedical Sciences, Heritage College of Osteopathic Medicine, Ohio University , Athens, OH 45701 , USA
4. Consejo Nacional de Investigaciones Científicas y Técnicas, División Paleontología Vertebrados, Museo de La Plata, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata , B1900 La Plata, Buenos Aires , Argentina
5. Evolutionary Studies Institute, University of the Witwatersrand , 2000 Johannesburg , South Africa
6. Department of Anatomical Sciences, Stony Brook University , Stony Brook, NY 11794 , USA