1. School of Fisheries, Aquaculture and Aquatic Sciences, College of Agriculture, Auburn University , Auburn, AL 36849 , USA
2. Southeast Conservation Genetics Laboratory, Warm Springs Fish Technology Center, United States Fish and Wildlife Service , Auburn, AL 36849 , USA
3. U.S. Geological Survey, Wetland and Aquatic Research Center , Gainesville, FL 32653 , USA
4. North Carolina Wildlife Resources Commission , Marion, NC 28652 , USA
5. United States Fish and Wildlife Service , Asheville, NC 28801 , USA