1. Department of Biological Sciences, University of Cyprus , Nicosia 2109 , Cyprus
2. Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences , Makhachkala 367000 , Russia
3. Department of Ecology, Dagestan State University , Makhachkala 367000 , Russia
4. Zoological Institute of the Russian Academy of Sciences , Saint Petersburg 199034 , Russia
5. Komarov Botanical Institute of the Russian Academy of Sciences , Saint Petersburg 197022 , Russia
6. Zoology Department, Section of Biology, Faculty of Science, Ege University , Bornova-Izmir 35100 , Turkey