1. Institute for Communities and Wildlife in Africa, Department of Biological Sciences, University of Cape Town , Cape Town , South Africa
2. Department of Botany and Zoology, Stellenbosch University , Stellenbosch , South Africa
3. Department of Zoology and Entomology, University of Pretoria , Pretoria , South Africa
4. Mpumalanga Tourism and Parks Agency , Lydenburg , South Africa
5. The Natural History Museum , London , United Kingdom
6. Institute for Biochemistry and Biology, University of Potsdam , Potsdam , Germany
7. Department of Zoology, University of Cambridge , Cambridge , United Kingdom