1. Ecology, Evolution, and Development Group, Department of Wetland Ecology, Estación Biológica de Doñana (CSIC) , Sevilla , Spain
2. Section of Herpetology, National Museums of Kenya , Nairobi 40658-00100 , Kenya
3. Zentralmagazin Naturwissenschaftlicher Sammlungen, Martin-Luther-Universität Halle-Wittenberg , Halle (Saale ), Germany
4. Life Sciences Department, Natural History Museum , London , UK