1. School of Biological Sciences, Faculty of Environmental and Life Sciences, University of Southampton , Southampton , UK
2. Bird Group, Natural History Museum , Akeman Street, Tring HP23 6AP , UK
3. Fossil Reptiles, Amphibians, and Birds Section, Natural History Museum , Cromwell Road, London SW7 5BD , UK
4. Oxford University Museum of Natural History , Parks Road, Oxford OX1 3PW , UK
5. Bioengineering Research Group, Faculty of Engineering and Physical Sciences, University of Southampton , Southampton , UK
6. Centre for Sport, Exercise and Osteoarthritis Research Versus Arthritis , Southampton , UK
7. Institute for Life Sciences, University of Southampton , Southampton , UK