1. Department of Zoology, Denver Museum of Nature & Science , 2001 Colorado Boulevard, Denver, CO 80205 , USA
2. Colorado State University , 711 Oval Drive, Fort Collins, CO 80521 , USA
3. Centro de Investigaciones Biológicas del Noroeste , Km 1, Carretera A, San Juan de La Costa, El Comitan, 23205 La Paz, BCS , Mexico