1. Department of Marine Organism Taxonomy & Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China
2. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, China
3. University of Chinese Academy of Sciences, Beijing, China
4. Laboratory for Marine Biology and Biotechnology, Pilot National Laboratory for Marine Science and Technology (Qingdao), Qingdao, China