1. ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, QLD, Australia
2. AIMS@JCU, James Cook University, Townsville, QLD, Australia
3. Australian Institute of Marine Science, PMB3, Townsville, QLD, Australia
4. Department of Invertebrate Zoology & Geology, California Academy of Sciences, San Francisco, CA, USA
5. Department of Biology, University of Nevada, Reno, Reno, NV, USA
6. Department of Biological Sciences, California State Polytechnic University, Pomona, West Temple Avenue, Pomona, CA, USA