1. Division of Ecology and Evolution, and Centre for Biodiversity Analysis, Research School of Biology, Australian National University, Canberra, ACT, Australia
2. Department of Ecology and Evolutionary Biology; Biodiversity Institute, University of Kansas, Lawrence, KS, USA
3. U.S. Geological Survey, Suite, San Diego, CA, USA
4. Aquatic Research Group, School of Civil Engineering, University of Queensland, Brisbane, QLD, Australia