1. Department of Marine Zoology-Crustaceans, Senckenberg Research Institute and Natural History Museum, Frankfurt am Main, Germany
2. Department of Ecology and Evolutionary Biology; and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS, USA
3. Florida Museum of Natural History, University of Florida, Gainesville, FL, USA
4. Smithsonian Tropical Research Institute, Balboa, Ancon, Republica de Panamá