1. Biodiversity, Ecology and Evolution Department, Faculty of Biology, Universidad Complutense de Madrid , 28040 Madrid , Spain
2. Grupo de Ecoloxía Animal (GEA), Universidade de Vigo , E-36310 Vigo , Spain
3. Eco&Sols, INRAE, IRD, CIRAD, SupAgro Montpellier , 34060 Montpellier , France
4. CEFE, Université de Montpellier, CNRS, EPHE, IRD , 34060 Montpellier , France