1. State Key Laboratory of Biocontrol, School of Life Sciences, Sun Yat-sen University , Guangzhou 510275 , China
2. GMU-GIBH Joint School of Life Sciences, Guangzhou Medical University , Guangzhou 511436 , China
3. Evolutionary Genomics and Modelling Lab, Centre for Evolution and Cancer, The Institute of Cancer Research , London SW7 3RP , UK
4. Department of Colorectal Surgery, The Sixth Affiliated Hospital, Sun Yat-sen University , Guangzhou 510655 , China
5. Guangdong Provincial Key Laboratory of Colorectal and Pelvic Floor Diseases, The Sixth Affiliated Hospital, Sun Yat-sen University , Guangzhou 510655 , China
6. Cancer Center, Clifford Hospital, Jinan University , Guangzhou 511495 , China