1. Beef , Sertãozinho, São Paulo 14174-000 , Brazil
2. Cattle Research Center, Animal Science Institute/APTA/SAA , Sertãozinho, São Paulo 14174-000 , Brazil
3. Department of Genetics, Ribeirão Preto Medical School, University of São Paulo , Ribeirão Preto, São Paulo, 14049-900 , Brazil
4. Centre for Genetic Improvement of Livestock, Department of Animal Biosciences, University of Guelph , Guelph, ON , Canada N1G 2W1