1. Behavioral Ecology Research Group, Center for Natural Sciences, University of Pannonia, Veszprém H-8200, Hungary
2. MTA-PE Evolutionary Ecology Research Group, University of Pannonia, Veszprém H-8200, Hungary
3. MTA-ELTE Comparative Ethology Research Group, Budapest, Hungary
4. Department of Biology, Theoretical Population Ecology and Evolution Group, Lund University, Lund, Sweden