1. Shaanxi Key Laboratory for Animal Conservation, Shaanxi Institute of Zoology, Xi’an, 710032, China
2. College of Life Sciences, Northwest University, Xi’an, 710069, China
3. Graduate School of Management, University of California Davis, Davis, CA 95616, USA
4. Beijing Zoo, Beijing, 100044, China
5. Center for Excellence in Animal Evolution and Genetics, Chinese Academy of Sciences, Kunming, 650223, China