1. Centre for Biodiversity & Environment Research, University College London, Gower Street, London, WC1E 6BT, UK
2. Dipartimento di Biologia, Università degli Studi di Firenze, Via Madonna del Piano, Sesto Fiorentino, Firenze, 50019, Italy
3. Department of Life Sciences and Systems Biology, University of Turin, Via Accademia Albertina 13, Turin, 10123, Italy
4. Department of Evolutionary Biology and Ecology, Institute of Biology I (Zoology), University of Freiburg, Hauptstraße 1, Freiburg (Brsg.), 79104, Germany