1. Laboratoire d’Ecologie, Systématique et Evolution, CNRS UMR 8079, Université-Saclay, Bâtiment 362, Orsay Cedex F-91405, France
2. Ministry of Education Key Laboratory for Biodiversity and Ecological Engineering, College of Life Sciences, Beijing Normal University, Beijing 100875, China
3. Zhejiang Tiantong Forest Ecosystem National Observation and Research Station, School of Ecological and Environmental Sciences, East China Normal University, Shanghai 200241, China