1. School of Earth and Ocean Sciences, University of Victoria , Victoria, BC V8P 5C2 , Canada
2. Met Office , Exeter EX1 3PB , UK
3. Invertebrate Paleontology, Academy of Natural Sciences of Drexel University , Philadelphia, PA 19103 , USA
4. Institute of Ocean Sciences, Fisheries and Oceans Canada , Sidney, BC V8L 4B2 , Canada
5. Department of Biology, University of Victoria , Victoria, BC V8P 5C2 , Canada