1. Department of Mathematics and Statistics, University of Strathclyde , 26 Richmond Street, Glasgow G1 1XH , UK
2. Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow , Graham Kerr Building, Glasgow G12 8QQ , UK
3. The Marine Biological Association, The Laboratory , Citadel Hill, Plymouth PL1 2PB , UK
4. Marine Scotland Science , 375 Victoria Road, Aberdeen AB11 9DB , UK