1. UMR 212 EME, IRD-IFREMER-UM2, Centre de Recherche Halieutique Méditerranéenne et Tropicale, Avenue Jean Monnet, BP 171, 34 200 Sète, France
2. Southwest Fisheries Science Center, NOAA Fisheries, 8604 La Jolla Shores Drive, La Jolla, CA 92037, USA
3. Pacific Islands Fisheries Science Center, NOAA Fisheries, 2570 Dole Street, Honolulu, Hawaii 96822-2396, USA
4. Département Ecologie et Modèles pour l'Halieutique, IFREMER, BP 21105, 44311 Nantes Cedex 03, France
5. Earth to Ocean Research Group, Department of Biological Sciences, Simon Fraser University, Burnaby, BC, CanadaV5A 1S6