1. Department of Fisheries and Wildlife, Oregon State University, Corvallis, OR 97331, USA
2. Department of Botany and Plant Pathology, Oregon State University, Corvallis, OR 97331, USA
3. Northwest Fisheries Science Center, NOAA, Hatfield Marine Science Center, Newport, OR 97365, USA
4. NOAA Earth System Research Laboratory, 325 Broadway, Boulder, CO 80305, USA